Ankarana Special Reserve in northern Madagascar was created in 1956.
It is a small, partially vegetated plateau composed of 150-million-year-old middle Jurassic limestone.
With an average annual rainfall of about 2 m (79 in), the underlying rocks are susceptible to erosion, thereby producing caves and underground rivers—a karst topography.
The rugged relief and the dense vegetation have helped protect the region from human intrusion.
The plateau slopes gently to the east, but on the west it ends abrubtly in the "Wall of Ankarana", a sheer cliff that extends 25 kilometres (16 mi) north to south, and rises as high as 280 metres (920 ft).
To the south, the limestone mass breaks up into separate spires known as tower karst.
In the center of the plateau, seismic activity and eons of rainfall have dissolved the limestone away in deep gorges, and sometimes redeposited it in ribbons of flowstone.
In places where the calcific upper layers have been completely eroded, the harder base rock has been etched into channels and ridges known as tsingy.
The largest sinkhole in the Ankarana karst region, Mangily sinkhole, measures up to 700 meters across and 140 meters deep with a volume of 25 million cubic meters.
The area is sacred to the Antankarana people, who have historically taken refuge from encroaching enemy armies in its caves and other natural rock shelters.
Reference: Wikipedia